Subgroups of the order Coleoptera

Subgroups of the order Coleoptera
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This article classifies the subgroups of the order Coleoptera (beetles) down to the level of families, following the system in Volume 2 of American Beetles (2002) for all North American families (there were changes in the system between Vols. 1 & 2), and following Lawrence & Newton (1995) for extralimital taxa. A few subfamilies, tribes and synonyms are shown here where a family has been reclassified.

Order Coleoptera (beetles)

Suborder Adephaga Schellenberg, 1806
- Amphizoidae LeConte, 1853
- Carabidae Latreille, 1802 (ground beetles)
  - Cicindelinae, formerly Cicindelidae Latreille, 1802 (tiger beetles)
  - Paussinae, formerly Paussidae Latreille, 1807
- Dytiscidae Leach, 1815
  
  Dytiscus latissimus, in the Dytiscidae
- Gyrinidae Latreille, 1802 (whirligig beetles)
- Haliplidae Aube, 1836
- Hygrobiidae Régimbart, 1878
- Noteridae C.G. Thomson, 1860
- Rhysodidae Laporte, 1840
- Trachypachidae C.G. Thomson, 1857
Suborder Archostemata Kolbe, 1908
- Crowsoniellidae Iablokoff-Khnzorian, 1983
- Cupedidae Laporte, 1836
- Jurodidae Ponomarenko, 1985
- Micromalthidae Barber, 1983 (telephone-pole beetle)
- Ommatidae Sharp and Muir, 1912
Suborder Myxophaga Crowson, 1955
- Hydroscaphidae LeConte, 1874
- Lepiceridae Hinton, 1936
- Sphaeriusidae Erichson, 1845 (=Microsporidae: Bulletin of Zoological Nomenclature 57(3):, 182-184.)
- Torridincolidae Steffan, 1964
Suborder Polyphaga Emery, 1886
- Infraorder Bostrichiformia Forbes, 1926
  - Superfamily Bostrichoidea Latreille, 1802
    - Anobiidae Fleming, 1821
      - Ptininae, formerly Ptinidae Latreille, 1802
    - Bostrichidae Latreille, 1802
      - Lyctinae, formerly Lyctidae Billberg, 1820
      - Endecatominae, formerly Endecatomidae LeConte, 1861
    - Dermestidae Latreille, 1804
      
      Anthrenus verbasci, in the Dermestidae
      - Thorictinae, formerly Thorictidae Agassiz, 1846
    - Jacobsoniidae Heller, 1926
    - Nosodendridae Erichson, 1846
  - Superfamily Derodontoidea LeConte, 1861
    - Derodontidae LeConte, 1861
- Infraorder Cucujiformia Lameere, 1938
  - Superfamily Chrysomeloidea Latreille, 1802
    - Cerambycidae Latreille, 1802 (longhorn beetles)
    - Chrysomelidae Latreille, 1802 (leaf beetles) - includes Bruchidae and Cassidae
      
      Colorado potato beetle, Leptinotarsa decemlineata, a leaf beetle
    - Megalopodidae Latreille, 1802
    - Orsodacnidae C. G. Thomson, 1869
  - Superfamily Cleroidea Latreille, 1802
    - Acanthocnemidae Crowson, 1964
    - Chaetosomatidae Crowson, 1952
    - Cleridae Latreille, 1802
    - Melyridae Leach, 1815
    - Phloiophilidae Kiesenwetter, 1863
    - Phycosecidae Crowson, 1952
    - Prionoceridae Lacordaire, 1857
    - Trogossitidae Latreille, 1802
  - Superfamily Cucujoidea Latreille, 1802
    - Alexiidae Imhoff, 1856
    - Biphyllidae LeConte, 1861
    - Boganiidae Sen Gupta and Crowson, 1966
    - Bothrideridae Erichson, 1845
    - Byturidae Jacquelin du Val, 1858
    - Cavognathidae Sen Gupta and Crowson, 1966
    - Cerylonidae Billberg, 1820
    - Coccinellidae Latreille, 1807 (ladybirds or lady beetles)
      
      Asian multicolored lady beetle, Harmonia axyridis
    - Corylophidae LeConte, 1852
    - Cryptophagidae Kirby, 1937
    - Cucujidae Latreille, 1802
    - Discolomatidae Horn, 1878
    - Endomychidae Leach, 1815
      - Merophysiinae, formerly Merophysiidae Seidlitz, 1872
    - Erotylidae Latreille, 1802
    - Helotidae Reitter, 1876
    - Hobartiidae Sen Gupta and Crowson, 1966
    - Kateretidae Erichson in Agassiz, 1846 (=Brachypteridae. ICZN Op., 1916, 1999).
    - Laemophloeidae Ganglbauer, 1899
    - Lamingtoniidae Sen Gupta and Crowson, 1966
    - Languriidae Crotch , 1873
    - Latridiidae Erichson, 1842
    - Monotomidae Laporte, 1840
      - Rhizophaginae, formerly Rhizophagidae Redtenbacher, 1845
    - Nitidulidae Latreille, 1802
    - Passandridae Erichson, 1845
    - Phalacridae Leach, 1815
    - Phloeostichidae Reitter, 1911
    - Propalticidae Crowson, 1952
    - Protocucujidae Crowson, 1954
    - Silvanidae Kirby, 1937
    - Smicripidae Horn, 1879
    - Sphindidae Jacquelin du Val, 1860
  - Superfamily Curculionoidea Latreille, 1802
    - Anthribidae Billberg, 1820
    - Attelabidae Billberg, 1820
    - Belidae Schönherr, 1826
      - Aglycyderinae, formerly Aglycyderidae Wollaston, 1864
      - Oxycoryninae, formerly Oxycorynidae Schönherr, 1840
    - Brentidae Billberg, 1820
      - Apioninae, formerly Apionidae Schönherr, 1823
    - Caridae Thompson, 1992
    - Curculionidae Latreille, 1802 (snout beetles and true weevils)
      
      Citrus root weevil, Diaprepes abbreviatus
      - Scolytinae, formerly Scolytidae Latreille, 1807
    - Ithyceridae Schönherr, 1823 (New York weevil)
    - Nemonychidae Bedel, 1882
  - Superfamily Lymexyloidea Fleming, 1821
    - Lymexylidae Fleming, 1821
  - Superfamily Tenebrionoidea Latreille, 1802
    - Aderidae Winkler, 1927
    - Anthicidae Latreille, 1819
    - Archeocrypticidae Kaszab, 1964
    - Boridae C. G. Thomson, 1859
    - Chalcodryidae Watt, 1974
    - Ciidae Leach, 1819 (=Cisidae)
    - Melandryidae Leach, 1815
    - Meloidae Gyllenhal, 1810 (blister beetles)
      
      Lytta magister, in the Meloidae
    - Mordellidae Latreille, 1802
    - Mycetophagidae Leach, 1815
    - Mycteridae Blanchard, 1845 (palm beetles and flower beetles)
      - Hemipeplinae, formerly Hemipeplidae Lacordaire, 1854
    - Oedemeridae Latreille, 1810
    - Perimylopidae St. George, 1939
    - Prostomidae C. G. Thomson, 1859
    - Pterogeniidae Crowson, 1953
    - Pyrochroidae Latreille, 1807
      - Cononotini or Cononotidae
      - Pedilinae, formerly Pedilidae Lacordaire, 1859
    - Pythidae Solier, 1834
    - Ripiphoridae Gemminger and Harold, 1870 formerly Rhipiphoridae
    - Salpingidae Leach, 1815
      - Elacatini or Elacatidae
      - Inopeplinae, formerly Inopeplidae Grouvelle, 1908
    - Scraptiidae Mulsant, 1856
    - Stenotrachelidae C. G. Thomson, 1859
      - Cephaloinae, formerly Cephaloidae LeConte, 1852
    - Synchroidae Lacordaire, 1859
    - Tenebrionidae Latreille, 1802 (darkling beetles)
      
      Heliotautus ruficollis, a darkling beetle
      - Alleculinae, formerly Alleculidae Laporte, 1840
      - Lagriinae, formerly Lagriidae Latreille, 1825
      - Nilionini or Nilionidae Lacordaire, 1859
      - Petriini or Petriidae
    - Tetratomidae Billberg, 1820
    - Trachelostenidae Lacordaire, 1859
    - Trictenotomidae Blanchard, 1845
    - Ulodidae Pascoe, 1869
    - Zopheridae Solier, 1834
      - Colydiinae or Colydiini, formerly Colydiidae Erichson, 1842
      - Monommatinae or Monommatini, formerly Monommatidae or Monommidae Blanchard, 1845
- Infraorder Elateriformia Crowson, 1960
  - Superfamily Buprestoidea Leach, 1815
    - Buprestidae Leach, 1815
    - Schizopodidae LeConte, 1861
  - Superfamily Byrrhoidea Latreille, 1804
    - Byrrhidae Latreille, 1804
    - Callirhipidae Emden, 1924
    - Chelonariidae Blanchard, 1845
    - Cneoglossidae Champion, 1897
    - Dryopidae Billberg, 1820
    - Elmidae Curtis, 1830
    - Eulichadidae Crowson, 1973
    - Heteroceridae MacLeay, 1825
    - Limnichidae Erichson, 1846
    - Lutrochidae Kasap and Crowson, 1975
    - Psephenidae Lacordaire, 1854
    - Ptilodactylidae Laporte, 1836
  - Superfamily Dascilloidea Guerin-Meneville, 1843
    - Dascillidae Guérin-Méneville, 1843 (1834)
    - Rhipiceridae Latreille, 1834
  - Superfamily Elateroidea Leach, 1815
    - Artematopodidae Lacordaire, 1857 (=Eurypogonidae)
    - Brachypsectridae Leconte and Horn, 1883
    - Cantharidae Imhoff, 1856
    - Cerophytidae Latreille, 1834
    - Drilidae Blanchard, 1845
    - Elateridae Leach, 1815 (click beetles)
      
      Alaus murina, a click beetle
      - Cebrioninae, formerly Cebrionidae Latreille, 1802
    - Eucnemidae Eschscholtz, 1829
    - Lampyridae Latreille, 1817 (fireflies)
    - Lycidae Laporte, 1836
    - Omalisidae Lacordaire, 1857
    - Omethidae LeConte, 1861
    - Phengodidae LeConte, 1861 (glowworms)
    - Plastoceridae Crowson, 1972
    - Podabrocephalidae Pic, 1930
    - Rhinorhipidae Lawrence, 1988
    - Telegeusidae Leng, 1920
    - Throscidae Laporte, 1840 (=Trixagidae)
  - Superfamily Scirtoidea Fleming, 1821
    - Clambidae Fischer, 1821
    - Decliniidae Nikitsky et al, 1994
    - Eucinetidae Lacordaire, 1857
    - Scirtidae Fleming, 1821 (=Helodidae)
- Infraorder Scarabaeiformia Crowson, 1960
  - Superfamily Scarabaeoidea Latreille, 1802
    - Belohinidae Paulian, 1959
    - Bolboceratidae Laporte de Castelnau, 1840
    - Ceratocanthidae White, 1842 (= Acanthoceridae)
    - Diphyllostomatidae Holloway, 1972
    - Geotrupidae Latreille, 1802
    - Glaphyridae MacLeay, 1819
    - Glaresidae Semenov-Tian-Shanskii and Medvedev, 1932
    - Hybosoridae Erichson, 1847
    - Lucanidae Latreille, 1804 (stag beetles)
      
      Lucanus cervus, a stag beetle
    - Ochodaeidae Mulsant and Rey, 1871
    - Passalidae Leach, 1815
    - Pleocomidae LeConte, 1861
    - Scarabaeidae Latreille, 1802
      - Dynastinae, formerly Dynastidae MacLeay, 1819 (rhinoceros beetles)
    - Trogidae MacLeay, 1819
- Infraorder Staphyliniformia Lameere, 1900
  - Superfamily Hydrophiloidea Latreille, 1802
    - Histeridae Gyllenhal, 1808
    - Hydrophilidae Latreille, 1802
      - Georyssinae, formerly Georyssidae Laporte, 1840
    - Sphaeritidae Schuckard, 1839
    - Synteliidae Lewis, 1882
  - Superfamily Staphylinoidea Latreille, 1802
    - Agyrtidae C.G. Thomson, 1859
    - Hydraenidae Mulsant, 1844
    - Leiodidae Fleming, 1821 (=Anisotomidae)
      - Platypsyllinae Ritsema, 1869 or Leptinidae
    - Ptiliidae Erichson, 1845
      
      Devil's coach horse beetle, Ocypus olens, a rove beetle
      - Cephaloplectinae, formerly Limulodidae Sharp, 1883
    - Scydmaenidae Leach, 1815
    - Silphidae Latreille, 1807
    - Staphylinidae Latreille, 1802 (rove beetles)
      - Scaphidiinae, formerly Scaphidiidae Latreille, 1807
      - Pselaphinae, formerly Pselaphidae Latreille, 1802

References

Lawrence, J.F., Newton, A.F. Jr. (1995) Families and subfamilies of Coleoptera (with selected genera, notes, references, and data on family-group names), pp. 779-1006. In: Pakaluk, J., Slipinski, S.A. (eds.), Biology, phylogeny, and classification of Coleoptera: Papers celebrating the 80th birthday of Roy A. Crowson. Muzeum i Instytut Zoologii PAN, Warszawa.
Ross H. Arnett, Jr. and Michael C. Thomas, American Beetles (CRC Press, 2001–2002)

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