The Pieridae are a large family of butterflies with about 76 genera containing approximately 1100 species, mostly from tropical Africa and Asia.[1] These butterflies are characterised by mainly white or yellow coloration, often with black spots. The sexes usually differ, often in the pattern or number of the black markings.
The Pieridae have the radial vein on the forewing with 3 or 4 branches and rarely with 5 branches. The fore legs are well developed in both sexes, unlike in the Nymphalidae, and the tarsal claws are bifid unlike in the Papilionidae.[2]
Like the Papilionidae, Pieridae also have their pupae held at an angle by a silk girdle, but running at the first abdominal segment unlike the thoracic girdle seen in the Papilionidae.
Subfamilies
The Pieridae are generally divided into the following four subfamilies:
Dismorphiinae (6 genera) Mostly Neotropical, this group includes several mimetical species. The host plants are in the family Fabaceae.[1]
Coliadinae (14 genera) (Sulphurs or Yellows, many of these species are sexually dimorphic. Some, such as Colias, have wing patterns that are visible only under ultraviolet.[1]
Pseudopontiinae The sole species in this subfamily, Pseudopontia paradoxa, is endemic to West Africa.
According to the molecular phylogenetic study of Braby et al. (2006), sister group relationships among Pieridae subfamilies are: ((Dismorphiinae+Pseudopontiinae)+(Coliadinae+Pierinae)).
^ abcde DeVries P. J. in Levin S.A. (ed) 2001 The Encyclopaedia of Biodiversity. Academic Press.
^ Borror, D. J., Triplehorn, C. A., & Johnson, N. F. (1989). An introduction to the study of insects (6th ed.). Philadelphia: Saunders College Publishers. ISBN 0030253977
Braby, M. F. 2005. Provisional checklist of genera of the Pieridae (Lepidoptera: Papilionidae). Zootaxa 832: 1–16.
Braby, M., R. Vila, and N. E. Pierce. 2006. Molecular phylogeny and systematics of the Pieridae (Lepidoptera: Papilionoidea: higher classification and biogeography. Zoological Journal of the Linnean Society 147(2): 239-275.