Ornithischia (pronounced /ɔrn
Characteristics
Edmontosaurus pelvis (showing ornithischian structure - left side) Oxford University Museum of Natural History
The Dinosauria superorder was divided into the two orders Ornithischia and Saurischia by Harry Seeley in 1887. This division, which has generally been accepted, is based on the evolution of the pelvis into a more bird-like structure (although birds did not descend from these dinosaurs), details in the vertebrae and armor and the possession of a 'predentary' bone. The predentary is an extra bone in the front of the lower jaw, which extends the dentary (the main lower jaw bone). The predentary coincides with the premaxilla in the upper jaw. Together they form a beak-like apparatus used to clip off plant material. The ornithischian pubis bone points downward and toward the tail (backwards), parallel with the ischium, with a forward-pointing process to support the abdomen. This makes a four-pronged pelvic structure. In contrast to this, the saurischian pubis points downward and towards the head (forwards), as in ancestral lizard types. Ornithischians also had smaller holes in front of their eye sockets (antorbital fenestrae) than saurischians, and a wider, more stable pelvis. A bird-like pubis arrangement, parallel to the vertebral column, independently evolved three times in dinosaur evolution, namely in the ornithischians, the therizinosauroids and in bird-like dromaeosaurids. ClassificationTaxonomyLinnaean ranks after Benton (2004),
PhylogenyThe ornithischians are divided in the two clades: the first are the Thyreophora and the second the Cerapoda. The Thyreophora include the Stegosauria (like the armored Stegosaurus) and the Ankylosauria (like Ankylosaurus). The Cerapoda include the Marginocephalia (Ceratopsia like the frilled ceratopsidae and Pachycephalosauria) and the Ornithopoda (among which duck-bills (hadrosaurs) such as Edmontosaurus). The Cerapoda are a relatively recent grouping (Sereno, 1986), and may conceivably be identical to (synonymous with) the older group, Ornithopoda: most of these divisions are not true by definition. Ornithischia |-?Pisanosaurus `--+-?Fabrosauridae `--Genasauria |--Thyreophora | |--Scutellosaurus | `--Thyreophoroidea | |--Emausaurus | `--Eurypoda | |--Stegosauria | `--Ankylosauromorpha | |--Scelidosaurus | `--Ankylosauria `--Cerapoda |--Stormbergia |--Agilisaurus |--Hexinlusaurus |--Heterodontosauridae `--+--Ornithopoda `--Marginocephalia |--Pachycephalosauria `--Ceratopsia (basal Cerapoda after Butler, 2005) References
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