Archimylacris lived on the warm swampyforest floors of North America and Europe300 million years ago, in the Late Carboniferous times. Like modern cockroaches, this insect had a large head shield with long curved antennae, or feelers, and folded wings. To a modern observer, it would likely appear as a giant cockroach with a "tail" (an ovipositor) in the females. Presumably their habits would be cockroach-like too, scurrying along the undergrowth eating anything edible, possibly falling prey to amphibians and very early reptiles.